About the Authors

Toshifumi Wada

wada@hitohaku.jp

Affiliation Institute of Natural and Environmental Sciences, University of Hyogo, Sanda, Hyogo, Japan

Takahiro Mitsushio

Affiliation Fukuoka Prefectural Fisheries High School, Fukutsu, Fukuoka, Japan

Shinya Inoue

Affiliation Kyushu Technological University, Kitakyushu, Fukuoka, Japan

Hiroko Koike

Affiliation Graduate School of Social and Cultural Studies, Kyushu University, Fukuoka, Japan

Ryo Kawabe

Affiliation Institute for East China Sea Research, Graduate School of Fisheries and Environmental Sciences, Nagasaki University, Nagasaki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TW RK. Performed the experiments: TW TM. Analyzed the data: TW SI RK. Contributed reagents/materials/analysis tools: TW HK RK. Wrote the paper: TW RK.