About the Authors

Qiaoqing Xu

Affiliations School of Animal Science, Yangtze University, Jingzhou, Hubei Province 434025, P. R. China, Scottish Fish Immunology Research Centre, Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 2TZ, United Kingdom, State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, Hubei Province 430072, P. R. China

Yousheng Jiang

Affiliations Scottish Fish Immunology Research Centre, Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 2TZ, United Kingdom, College of Fishery and Life Science, Shanghai Ocean University, Shanghai 201306, China

Eakapol Wangkahart

Affiliation Scottish Fish Immunology Research Centre, Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 2TZ, United Kingdom

Jun Zou

Affiliation Scottish Fish Immunology Research Centre, Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 2TZ, United Kingdom

Mingxian Chang

Affiliation State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, Hubei Province 430072, P. R. China

Daiqin Yang

Affiliation School of Animal Science, Yangtze University, Jingzhou, Hubei Province 434025, P. R. China

Chris J. Secombes

Affiliation Scottish Fish Immunology Research Centre, Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 2TZ, United Kingdom

Pin Nie

t.h.wang@abdn.ac.uk (TW); pinnie@ihb.ac.cn (PN)

Affiliation State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, Hubei Province 430072, P. R. China

Tiehui Wang

t.h.wang@abdn.ac.uk (TW); pinnie@ihb.ac.cn (PN)

Affiliation Scottish Fish Immunology Research Centre, Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 2TZ, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: QX PN TW. Performed the experiments: QX YJ EW JZ MC DY TW. Analyzed the data: QX TW. Wrote the paper: QX DY CJS PN TW.