About the Authors

Marie-Jose Naud

Affiliation School of Biological Sciences, Royal Holloway University of London, Egham, United Kingdom

Warwick H. H. Sauer

Affiliation Department of Ichthyology and Fisheries Science, Rhodes University, Grahamstown, South Africa

Niall J. McKeown

Affiliation Institute of Biological, Environmental & Rural Sciences, Aberystwyth University, Aberystwyth, United Kingdom

Paul W. Shaw

pws3@aber.ac.uk

Affiliations Institute of Biological, Environmental & Rural Sciences, Aberystwyth University, Aberystwyth, United Kingdom, Department of Ichthyology and Fisheries Science, Rhodes University, Grahamstown, South Africa

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: M-JN WHHS PWS. Performed the experiments: M-JN NJM PWS. Analyzed the data: M-JN NJM PWS. Contributed reagents/materials/analysis tools: PWS WHHS. Wrote the paper: M-JN WHHS NJM PWS. Fieldwork: WHHS PWS.