About the Authors

Minami Orii

Affiliation Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, 1 Kawasumi, Mizuho-cho, Mizuho-ku, Nagoya 467–8601, Japan

Keiko Kono

mkt-naka@med.nagoya-cu.ac.jp (MN); kono@med.nagoya-cu.ac.jp (KK)

Affiliation Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, 1 Kawasumi, Mizuho-cho, Mizuho-ku, Nagoya 467–8601, Japan

Hsin-I Wen

Affiliation Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, 1 Kawasumi, Mizuho-cho, Mizuho-ku, Nagoya 467–8601, Japan

Makoto Nakanishi

mkt-naka@med.nagoya-cu.ac.jp (MN); kono@med.nagoya-cu.ac.jp (KK)

Affiliation Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, 1 Kawasumi, Mizuho-cho, Mizuho-ku, Nagoya 467–8601, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MN KK. Performed the experiments: MO HW KK. Analyzed the data: MO HW KK. Contributed reagents/materials/analysis tools: MO HW KK. Wrote the paper: MN KK MO.