About the Authors

Hazuki Yamada

Affiliation Division of Cell Biology, Department of Biosciences, School of Science, Kitasato University, Kanagawa, 252–0373, Japan

Koji Tsutsumi

Affiliation Division of Cell Biology, Department of Biosciences, School of Science, Kitasato University, Kanagawa, 252–0373, Japan

Yuki Nakazawa

Affiliation Division of Cell Biology, Department of Biosciences, School of Science, Kitasato University, Kanagawa, 252–0373, Japan

Yoshio Shibagaki

Affiliation Division of Biochemistry, School of Pharmaceutical Sciences, Kitasato University, Tokyo, 108–8641, Japan

Seisuke Hattori

Affiliation Division of Biochemistry, School of Pharmaceutical Sciences, Kitasato University, Tokyo, 108–8641, Japan

Yasutaka Ohta

yohta@kitasato-u.ac.jp

Affiliation Division of Cell Biology, Department of Biosciences, School of Science, Kitasato University, Kanagawa, 252–0373, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HY KT YN YS SH YO. Performed the experiments: HY KT YN YS. Analyzed the data: HY KT YN YS SH YO. Contributed reagents/materials/analysis tools: HY KT YN YS. Wrote the paper: HY KT SH YO.