About the Authors

Victoria J. Brookes

victoria.brookes@sydney.edu.au

Affiliations Graham Centre for Agricultural Innovation, NSW Department of Primary Industries and Charles Sturt University, Pugsley Place, Wagga Wagga, NSW 2650, Australia, School of Animal and Veterinary Sciences, Faculty of Science, Charles Sturt University, Wagga Wagga, NSW 2678, Australia

David Jordan

Affiliation NSW Department of Primary Industries, Wollongbar, NSW, Australia

Stephen Davis

Affiliation School of Mathematical and Geospatial Sciences, RMIT University, Melbourne, Victoria 3001, Australia

Michael P. Ward

Affiliation Faculty of Veterinary Science, The University of Sydney, Camden, NSW 2570, Australia

Jane Heller

Affiliations Graham Centre for Agricultural Innovation, NSW Department of Primary Industries and Charles Sturt University, Pugsley Place, Wagga Wagga, NSW 2650, Australia, School of Animal and Veterinary Sciences, Faculty of Science, Charles Sturt University, Wagga Wagga, NSW 2678, Australia

Competing Interests

The authors have the following interests: This research was funded by Meat and Livestock Australia (MLA; A.MFS.0247). There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: VJB DJ SD MPW JH. Performed the experiments: VJB. Analyzed the data: VJB. Contributed reagents/materials/analysis tools: VJB DJ SD. Wrote the paper: VJB DJ SD MPW JH.