About the Authors

Zarinah Waheed

zarinah.waheed@naturalis.nl

Affiliations Department of Marine Zoology, Naturalis Biodiversity Center, Leiden, The Netherlands, Borneo Marine Research Institute, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia, Institute of Biology Leiden, Leiden University, Leiden, The Netherlands

Harald G. J. van Mil

Affiliation Institute of Biology Leiden, Leiden University, Leiden, The Netherlands

Muhammad Ali Syed Hussein

Affiliation Borneo Marine Research Institute, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia

Robecca Jumin

Affiliation WWF-Malaysia, Kota Kinabalu, Sabah, Malaysia

Bobita Golam Ahad

Affiliation WWF-Malaysia, Kota Kinabalu, Sabah, Malaysia

Bert W. Hoeksema

Affiliation Department of Marine Zoology, Naturalis Biodiversity Center, Leiden, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZW HGJVM MASH RJ BGA BWH. Performed the experiments: ZW MASH RJ BGA BWH. Analyzed the data: ZW HGJVM. Contributed reagents/materials/analysis tools: ZW HGJVM MASH RJ BGA BWH. Wrote the paper: ZW HGJVM MASH BWH.