About the Authors

Jani Anttila

jani.anttila@helsinki.fi

Affiliation Integrative Ecology Unit, Department of Biosciences, FI-00014 University of Helsinki, Helsinki, Finland

Veijo Kaitala

Affiliation Integrative Ecology Unit, Department of Biosciences, FI-00014 University of Helsinki, Helsinki, Finland

Jouni Laakso

Affiliation Integrative Ecology Unit, Department of Biosciences, FI-00014 University of Helsinki, Helsinki, Finland

Lasse Ruokolainen

Affiliation Integrative Ecology Unit, Department of Biosciences, FI-00014 University of Helsinki, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: JA JL VK LR. Provided the original idea: LR. Constructed the model: JA. Analysed the model: JA LR.