About the Authors

Johanna Rytioja

Affiliation Department of Food and Environmental Sciences, Division of Microbiology and Biotechnology, University of Helsinki, Helsinki, Finland

Kristiina Hildén

Affiliation Department of Food and Environmental Sciences, Division of Microbiology and Biotechnology, University of Helsinki, Helsinki, Finland

Susanna Mäkinen

Affiliation Roal Oy, Rajamäki, Finland

Jari Vehmaanperä

Affiliation Roal Oy, Rajamäki, Finland

Annele Hatakka

Affiliation Department of Food and Environmental Sciences, Division of Microbiology and Biotechnology, University of Helsinki, Helsinki, Finland

Miia R. Mäkelä

miia.r.makela@helsinki.fi

Affiliation Department of Food and Environmental Sciences, Division of Microbiology and Biotechnology, University of Helsinki, Helsinki, Finland

Competing Interests

The authors declare that they have no competing interests. The authors confirm that the fact that two of the authors of the manuscript are employed by a commercial company Roal Oy does not alter their adherence to PLOS ONE policies on sharing data and material.

Author Contributions

Conceived and designed the experiments: JR KH AH MRM. Performed the experiments: JR SM MRM. Analyzed the data: JR. Contributed reagents/materials/analysis tools: AH JV. Wrote the paper: JR KH SM JV AH MRM.