About the Authors

Huilin Gao

Affiliation Department of Forest Management, School of Forestry, Northeast Forestry University, Harbin, Heilongjiang, People’s Republic of China

Lihu Dong

Affiliation Department of Forest Management, School of Forestry, Northeast Forestry University, Harbin, Heilongjiang, People’s Republic of China

Fengri Li

fengrili@126.com

Affiliation Department of Forest Management, School of Forestry, Northeast Forestry University, Harbin, Heilongjiang, People’s Republic of China

Lianjun Zhang

Affiliation Department of Forest and Natural Resources Management, College of Environmental Science and Forestry, State University of New York, Old Westbury, NY, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FL HLG. Performed the experiments: HLG LHD. Analyzed the data: HLG. Contributed reagents/materials/analysis tools: HLG LHD FL LJZ. Wrote the paper: HLG. Provided detailed revised comments to the manuscript: LJZ LHD.