About the Authors

Megan E. Moriarty

Affiliation Karen C. Drayer Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

T. Winston Vickers

twvickers@ucdavis.edu (TWV); wmboyce@ucdavis.edu (WMB)

Affiliations Karen C. Drayer Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America, Institute for Wildlife Studies, Arcata, California, United States of America

Deana L. Clifford

Affiliations Karen C. Drayer Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America, Wildlife Investigations Laboratory, California Department of Fish and Wildlife, Rancho Cordova, California, United States of America

David K. Garcelon

Affiliation Institute for Wildlife Studies, Arcata, California, United States of America

Patricia M. Gaffney

Affiliations Department of Pathology, Microbiology, and Immunology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America, Departments of Pathology and Medicine, University of California San Diego, San Diego, California, United States of America

Kenneth W. Lee

Affiliation Greer Laboratory, Lenoir, North Carolina, United States of America

Julie L. King

Affiliation Catalina Island Conservancy, Avalon, California, United States of America

Calvin L. Duncan

Affiliation Catalina Island Conservancy, Avalon, California, United States of America

Walter M. Boyce

twvickers@ucdavis.edu (TWV); wmboyce@ucdavis.edu (WMB)

Affiliations Karen C. Drayer Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America, Department of Pathology, Microbiology, and Immunology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Competing Interests

We have the following interests: This study received funding from Greer Laboratories, Inc. Kenneth W. Lee is employed by Greer Laboratories, Inc. There are no patents, products in development, or marketed products as a result of this project. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MEM TWV DLC DKG KWL JLK CLD WMB. Performed the experiments: MEM TWV DLC PMG KWL JLK CLD WMB. Analyzed the data: MEM TWV DLC DKG PMG KWL JLK WMB. Contributed reagents/materials/analysis tools: DLC DKG PMG KWL JLK WMB. Wrote the paper: MEM TWV DLC DKG PMG KWL JLK CLD WMB.