About the Authors

Meagan C. Burnet

Affiliation Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, 99352, United States of America

Alice C. Dohnalkova

Affiliation Environmental Molecular Sciences Laboratory, Pacific Northwest National Laboratory, Richland, Washington, 99352, United States of America

Anthony P. Neumann

Affiliation Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, 53706, United States of America

Mary S. Lipton

Affiliation Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, 99352, United States of America

Richard D. Smith

Affiliation Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, 99352, United States of America

Garret Suen

stephen.callister@pnl.gov (SJC); gsuen@wisc.edu (GS)

Affiliation Department of Bacteriology, University of Wisconsin-Madison, Madison, Wisconsin, 53706, United States of America

Stephen J. Callister

stephen.callister@pnl.gov (SJC); gsuen@wisc.edu (GS)

Affiliation Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, 99352, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MCB ACD APN GS SJC. Performed the experiments: MCB ACD APN. Analyzed the data: MCB APN. Contributed reagents/materials/analysis tools: MCB ACD APN RDS GS. Wrote the paper: MCB ACD APN MSL SJC GS.