About the Authors

Russell W. Avramenko

Affiliation Department of Comparative Biology and Experimental Medicine, Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

Elizabeth M. Redman

Affiliation Department of Comparative Biology and Experimental Medicine, Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

Roy Lewis

Affiliation Merck Animal Health, Calgary, Alberta, Canada

Thomas A. Yazwinski

Affiliation Department of Animal Science, University of Arkansas, Fayetteville, Arkansas, United States of America

James D. Wasmuth

Affiliation Department of Ecosystem and Public Health, Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

John S. Gilleard

jsgillea@ucalgary.ca

Affiliation Department of Comparative Biology and Experimental Medicine, Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

Competing Interests

Roy Lewis is employed with Merck Animal Health as a technical Services Veterinarian part-time. This affiliation does not alter the authors' adherence to PLOS ONE policies on sharing data and materials. The other authors declare that no other competing interests exist.

Author Contributions

Conceived and designed the experiments: RWA JSG. Performed the experiments: RWA EMR. Analyzed the data: RWA. Contributed reagents/materials/analysis tools: JSG JDW RL TAY. Wrote the paper: RWA JSG.