About the Authors

T. Winston Vickers

twvickers@ucdavis.edu (TWV), wmboyce@ucdavis.edu (WMB)

Affiliations Institute for Wildlife Studies, Arcata, California, United States of America, Karen C. Drayer Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Deana L. Clifford

Affiliations Karen C. Drayer Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America, Wildlife Investigations Lab, California Department of Fish and Wildlife, Rancho Cordova, California, United States of America

David K. Garcelon

Affiliation Institute for Wildlife Studies, Arcata, California, United States of America

Julie L. King

Affiliation Catalina Island Conservancy, Avalon, California, United States of America

Calvin L. Duncan

Affiliation Catalina Island Conservancy, Avalon, California, United States of America

Patricia M. Gaffney

Affiliations Department of Pathology, Microbiology, and Immunology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America, Departments of Pathology and Medicine, University of California San Diego, San Diego, California, United States of America

Walter M. Boyce

twvickers@ucdavis.edu (TWV), wmboyce@ucdavis.edu (WMB)

Affiliations Karen C. Drayer Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America, Department of Pathology, Microbiology, and Immunology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TWV DLC DKG JLK CLD WMB. Performed the experiments: TWV DLC JLK CLD PMG WMB. Analyzed the data: TWV DLC DKG JLK PMG WMB. Contributed reagents/materials/analysis tools: DLC DKG JLK PMG WMB. Wrote the paper: TWV DLC DKG JLK CLD PMG WMB.