About the Authors

Shumpei Asamizu

asamizu@mail.ecc.u-tokyo.ac.jp (SA); aonaka@mail.ecc.u-tokyo.ac.jp (HO)

Affiliation Department of Biotechnology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo, Tokyo, Japan

Taro Ozaki

Affiliation Department of Biotechnology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo, Tokyo, Japan

Kanae Teramoto

Affiliation Advanced Technology Department, JEOL Ltd., Akishima, Tokyo, Japan

Katsuya Satoh

Affiliation Ion Beam Mutagenesis Research Group, Biotechnology and Medical Application Division, Quantum Beam Science Center, Japan Atomic Energy Agency, Takasaki, Gunma, Japan

Hiroyasu Onaka

asamizu@mail.ecc.u-tokyo.ac.jp (SA); aonaka@mail.ecc.u-tokyo.ac.jp (HO)

Affiliation Department of Biotechnology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo, Tokyo, Japan

Competing Interests

Kanae Teramoto is employed by Advanced Technology Department, JEOL Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: SA HO. Performed the experiments: SA KT. Analyzed the data: SA TO KT KS HO. Contributed reagents/materials/analysis tools: KT KS HO. Wrote the paper: SA HO. Revised the manuscript: TO.