About the Authors

Emi Kawano-Yamashita

Affiliation Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan

Mitsumasa Koyanagi

Affiliations Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan, The OCU Advanced Research Institute for Natural Science and Technology (OCARINA), Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan, Japan Science and Technology Agency (JST), Precursory Research for Embryonic Science and Technology (PRESTO), Tokyo, Japan

Seiji Wada

Affiliation Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan

Hisao Tsukamoto

Affiliation Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan

Takashi Nagata

Affiliation Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan

Akihisa Terakita

terakita@sci.osaka-cu.ac.jp

Affiliations Department of Biology and Geosciences, Graduate School of Science, Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan, The OCU Advanced Research Institute for Natural Science and Technology (OCARINA), Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka, 558–8585, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EK-Y MK AT. Performed the experiments: EK-Y MK SW HT TN. Analyzed the data: EK-Y MK SW HT TN AT. Contributed reagents/materials/analysis tools: AT. Wrote the paper: EK-Y MK AT.