About the Authors

Joshua F. Goldberg

joshua.goldberg@umontana.edu

Current address: Evolution, Ecology and Organismal Biology Program, Department of Biology, University of California Riverside, Riverside, California, United States of America

Affiliation Wildlife Biology Program, Department of Ecosystem and Conservation Science, College of Forestry and Conservation, University of Montana, Missoula, Montana, United States of America

Tshering Tempa

Affiliations Wildlife Biology Program, Department of Ecosystem and Conservation Science, College of Forestry and Conservation, University of Montana, Missoula, Montana, United States of America, Ugyen Wangchuck Institute for Conservation and Environment, Department of Forests and Park Services, Ministry of Agriculture and Forests, Lamai Goempa, Bumthang, Bhutan

Nawang Norbu

Affiliation Ugyen Wangchuck Institute for Conservation and Environment, Department of Forests and Park Services, Ministry of Agriculture and Forests, Lamai Goempa, Bumthang, Bhutan

Mark Hebblewhite

Affiliation Wildlife Biology Program, Department of Ecosystem and Conservation Science, College of Forestry and Conservation, University of Montana, Missoula, Montana, United States of America

L. Scott Mills

Affiliation Department of Forestry and Environmental Resources, North Carolina State University, Raleigh, North Carolina, United States of America

Tshewang R. Wangchuk

Affiliation Bhutan Foundation, Washington, D.C., United States of America

Paul Lukacs

Affiliation Wildlife Biology Program, Department of Ecosystem and Conservation Science, College of Forestry and Conservation, University of Montana, Missoula, Montana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JFG; TT; MH; LSM. Performed the experiments: TT. Analyzed the data: JFG; TT. Contributed reagents/materials/analysis tools: JFG; TT; NN; MH; LSM; TRW; PL. Wrote the paper: JFG; TT; MH; LSM; PL.