About the Authors

Jae Hoon Shin

Contributed equally to this work with: Jae Hoon Shin, Il Yong Kim

Affiliations Laboratory of Developmental Biology and Genomics, College of Veterinary Medicine, and BK21 Program for Veterinary Science, Seoul National University, Seoul, South Korea, Korea Mouse Phenotyping Center (KMPC), Seoul National University, Seoul, South Korea

Il Yong Kim

Contributed equally to this work with: Jae Hoon Shin, Il Yong Kim

Affiliations Laboratory of Developmental Biology and Genomics, College of Veterinary Medicine, and BK21 Program for Veterinary Science, Seoul National University, Seoul, South Korea, Korea Mouse Phenotyping Center (KMPC), Seoul National University, Seoul, South Korea

Yo Na Kim

Affiliations Laboratory of Developmental Biology and Genomics, College of Veterinary Medicine, and BK21 Program for Veterinary Science, Seoul National University, Seoul, South Korea, Korea Mouse Phenotyping Center (KMPC), Seoul National University, Seoul, South Korea

Sun Mee Shin

Affiliation Division of Life Sciences, Ewha Womans University, Seoul, South Korea

Kyung Jin Roh

Affiliation Lee Gil Ya Cancer and Diabetes Institute and Division of Endocrinology Gil Medical Center, Gachon University of Medicine and Science, Incheon, South Korea

Seo Hyun Lee

Affiliations Laboratory of Developmental Biology and Genomics, College of Veterinary Medicine, and BK21 Program for Veterinary Science, Seoul National University, Seoul, South Korea, Korea Mouse Phenotyping Center (KMPC), Seoul National University, Seoul, South Korea

Mira Sohn

Affiliation Division of Life Sciences, Ewha Womans University, Seoul, South Korea

Soo Young Cho

Affiliations Laboratory of Developmental Biology and Genomics, College of Veterinary Medicine, and BK21 Program for Veterinary Science, Seoul National University, Seoul, South Korea, Korea Mouse Phenotyping Center (KMPC), Seoul National University, Seoul, South Korea

Sang Hyuk Lee

Affiliation Ewha Research Center for Systems Biology, Ewha Womans University, Seoul, South Korea

Chang-Yong Ko

Affiliation Department of Biomedical Engineering, College of Health Science, Institute of Medical Engineering, Yonsei University, Wonju, South Korea

Han-Sung Kim

Affiliations Laboratory of Developmental Biology and Genomics, College of Veterinary Medicine, and BK21 Program for Veterinary Science, Seoul National University, Seoul, South Korea, Korea Mouse Phenotyping Center (KMPC), Seoul National University, Seoul, South Korea

Cheol Soo Choi

Affiliation Lee Gil Ya Cancer and Diabetes Institute and Division of Endocrinology Gil Medical Center, Gachon University of Medicine and Science, Incheon, South Korea

Yun Soo Bae

snumouse@snu.ac.kr (JKS); baeys@ewha.ac.kr (YSB)

Affiliation Division of Life Sciences, Ewha Womans University, Seoul, South Korea

Je Kyung Seong

snumouse@snu.ac.kr (JKS); baeys@ewha.ac.kr (YSB)

Affiliations Laboratory of Developmental Biology and Genomics, College of Veterinary Medicine, and BK21 Program for Veterinary Science, Seoul National University, Seoul, South Korea, Korea Mouse Phenotyping Center (KMPC), Seoul National University, Seoul, South Korea, Interdisciplinary Program for Bioinformatics, Program for Cancer Biology and BIO-MAX Institute, Seoul National University, Seoul, South Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JHS IYK JKS. Performed the experiments: JHS IYK YNK KJR Seo Hyun L. CYK. Analyzed the data: JHS YNK CYK CSC SYC Sang Hyuk L. Contributed reagents/materials/analysis tools: SMS MS HSK CSC YSB. Wrote the paper: JHS IYK JKS.