About the Authors

Shu Liao

Affiliations Long-term Research Station of Alpine Forest Ecosystems, Key Laboratory of Ecological Forestry Engineering in the Upper Reaches of the Yangtze River of Sichuan Province, Institute of Ecology and Forestry, Sichuan Agricultural University, Chengdu, China, Collaborative Innovation Center of Ecological Security in the Upper Reaches of the Yangtze River, Chengdu, China

Wanqin Yang

Affiliations Long-term Research Station of Alpine Forest Ecosystems, Key Laboratory of Ecological Forestry Engineering in the Upper Reaches of the Yangtze River of Sichuan Province, Institute of Ecology and Forestry, Sichuan Agricultural University, Chengdu, China, Collaborative Innovation Center of Ecological Security in the Upper Reaches of the Yangtze River, Chengdu, China

Yu Tan

Affiliations Long-term Research Station of Alpine Forest Ecosystems, Key Laboratory of Ecological Forestry Engineering in the Upper Reaches of the Yangtze River of Sichuan Province, Institute of Ecology and Forestry, Sichuan Agricultural University, Chengdu, China, Collaborative Innovation Center of Ecological Security in the Upper Reaches of the Yangtze River, Chengdu, China

Yan Peng

Affiliations Long-term Research Station of Alpine Forest Ecosystems, Key Laboratory of Ecological Forestry Engineering in the Upper Reaches of the Yangtze River of Sichuan Province, Institute of Ecology and Forestry, Sichuan Agricultural University, Chengdu, China, Collaborative Innovation Center of Ecological Security in the Upper Reaches of the Yangtze River, Chengdu, China

Jun Li

Affiliations Long-term Research Station of Alpine Forest Ecosystems, Key Laboratory of Ecological Forestry Engineering in the Upper Reaches of the Yangtze River of Sichuan Province, Institute of Ecology and Forestry, Sichuan Agricultural University, Chengdu, China, Collaborative Innovation Center of Ecological Security in the Upper Reaches of the Yangtze River, Chengdu, China

Bo Tan

Affiliations Long-term Research Station of Alpine Forest Ecosystems, Key Laboratory of Ecological Forestry Engineering in the Upper Reaches of the Yangtze River of Sichuan Province, Institute of Ecology and Forestry, Sichuan Agricultural University, Chengdu, China, Collaborative Innovation Center of Ecological Security in the Upper Reaches of the Yangtze River, Chengdu, China

Fuzhong Wu

wufzchina@163.com

Affiliations Long-term Research Station of Alpine Forest Ecosystems, Key Laboratory of Ecological Forestry Engineering in the Upper Reaches of the Yangtze River of Sichuan Province, Institute of Ecology and Forestry, Sichuan Agricultural University, Chengdu, China, Collaborative Innovation Center of Ecological Security in the Upper Reaches of the Yangtze River, Chengdu, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SL WY BT FW. Performed the experiments: SL YT YP JL. Analyzed the data: SL. Contributed reagents/materials/analysis tools: FW. Wrote the paper: SL FW.