About the Authors

Kyle Friend

friendk@wlu.edu (KF); jekimble@wisc.edu (JK)

Affiliations Department of Biochemistry, University of Wisconsin-Madison, Madison, Wisconsin, 53706, United States of America, Department of Chemistry and Biochemistry, Washington and Lee University, Lexington, Virginia, 24450, United States of America

Hunter A. Brooks

Affiliation Department of Chemistry and Biochemistry, Washington and Lee University, Lexington, Virginia, 24450, United States of America

Nicholas E. Propson

Affiliation The Morgridge Institute for Research, 309 North Orchard Street, Madison, Wisconsin, 53715, United States of America

James A. Thomson

Affiliations The Morgridge Institute for Research, 309 North Orchard Street, Madison, Wisconsin, 53715, United States of America, Department of Cell and Regenerative Biology, University of Wisconsin School of Medicine and Public Health, Madison, Wisconsin, 53706, United States of America

Judith Kimble

friendk@wlu.edu (KF); jekimble@wisc.edu (JK)

Affiliations Department of Biochemistry, University of Wisconsin-Madison, Madison, Wisconsin, 53706, United States of America, Howard Hughes Medical Institute, University of Wisconsin-Madison, Madison, Wisconsin, 53706, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KF JK. Performed the experiments: KF HAB NEP. Analyzed the data: KF. Contributed reagents/materials/analysis tools: NEP JAT. Wrote the paper: KF JAT JK.