About the Authors
- Naoto Tanaka
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Contributed equally to this work with: Naoto Tanaka, Emily V. Dutrow
Affiliation Department of Biology, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America
- Emily V. Dutrow
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Contributed equally to this work with: Naoto Tanaka, Emily V. Dutrow
Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America
- Keiko Miyadera
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Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America
- Lucie Delemotte
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Affiliation Institute for Computational Molecular Science, Temple University, Philadelphia, Pennsylvania, United States of America
- Christopher M. MacDermaid
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Affiliation Institute for Computational Molecular Science, Temple University, Philadelphia, Pennsylvania, United States of America
- Shelby L. Reinstein
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Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America
- William R. Crumley
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Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America
- Christopher J. Dixon
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Affiliation Veterinary Vision, Penrith, Cumbria, CA11 9FJ, United Kingdom
- Margret L. Casal
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Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America
- Michael L. Klein
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Affiliation Institute for Computational Molecular Science, Temple University, Philadelphia, Pennsylvania, United States of America
- Gustavo D. Aguirre
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Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America
- Jacqueline C. Tanaka
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* E-mail: jtanaka@temple.edu (JCT); karinag@vet.upenn.edu (KEG)
Affiliation Department of Biology, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America
- Karina E. Guziewicz
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* E-mail: jtanaka@temple.edu (JCT); karinag@vet.upenn.edu (KEG)
Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America
Competing Interests
CJD is a co-owner of a private veterinary practice, Veterinary Vision (Penrith, UK). This affiliation did not have any role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. This does not alter the authors' adherence to all PLOS ONE polices on sharing data and materials. All remaining authors declare no conflicts of interest.
Author Contributions
Conceived and designed the experiments: KEG GDA EVD JCT NT. Performed the experiments: EVD KM SLR WRC MLC KEG GDA JCT CMM LD NT. Analyzed the data: KEG EVD KM CJD JCT CMM GDA LD NT. Contributed reagents/materials/analysis tools: GDA KEG EVD KM CJD MLC JCT CMM LD NT. Wrote the paper: EVD NT KEG JCT CMM LD GDA MLK.