About the Authors

Yohei Nishikawa

Contributed equally to this work with: Yohei Nishikawa, Masahito Hosokawa

Affiliation Department of Life Science and Medical Bioscience, Waseda University, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Masahito Hosokawa

Contributed equally to this work with: Yohei Nishikawa, Masahito Hosokawa

Affiliations Department of Life Science and Medical Bioscience, Waseda University, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan, Research Organization for Nano & Life Innovation, Waseda University, 513, Wasedatsurumaki-cho, Shinjuku-ku, Tokyo 162–0041, Japan, Core Research for Evolutionary Science and Technology (CREST), Japan Science and Technology Agency (JST), 5, Sanbancho, Chiyoda-ku, Tokyo 102–0075, Japan

Toru Maruyama

Affiliation Department of Life Science and Medical Bioscience, Waseda University, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Keisuke Yamagishi

Affiliation Department of Life Science and Medical Bioscience, Waseda University, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan

Tetsushi Mori

Affiliations Department of Life Science and Medical Bioscience, Waseda University, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan, Core Research for Evolutionary Science and Technology (CREST), Japan Science and Technology Agency (JST), 5, Sanbancho, Chiyoda-ku, Tokyo 102–0075, Japan

Haruko Takeyama

haruko-takeyama@waseda.jp

Affiliations Department of Life Science and Medical Bioscience, Waseda University, 2–2 Wakamatsu-cho, Shinjuku-ku, Tokyo 162–8480, Japan, Research Organization for Nano & Life Innovation, Waseda University, 513, Wasedatsurumaki-cho, Shinjuku-ku, Tokyo 162–0041, Japan, Core Research for Evolutionary Science and Technology (CREST), Japan Science and Technology Agency (JST), 5, Sanbancho, Chiyoda-ku, Tokyo 102–0075, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN MH HT. Performed the experiments: YN KY T. Mori. Analyzed the data: YN MH T. Maruyama. Contributed reagents/materials/analysis tools: YN MH KY T. Mori. Wrote the paper: YN MH HT.