About the Authors

Megan Ulmer Carnes

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Terry Campbell

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Wen Huang

Affiliations Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America, Program in Genetics, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Daniel G. Butler

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Mary Anna Carbone

Affiliations Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America, Program in Genetics, North Carolina State University, Raleigh, North Carolina, 27695, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Laura H. Duncan

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Sasha V. Harbajan

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Edward M. King

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Kara R. Peterson

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Alexander Weitzel

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Shanshan Zhou

Affiliations Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America, Program in Genetics, North Carolina State University, Raleigh, North Carolina, 27695, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Trudy F. C. Mackay

trudy_mackay@ncsu.edu

Affiliations Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, 27695, United States of America, Program in Genetics, North Carolina State University, Raleigh, North Carolina, 27695, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, 27695, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TFCM TC. Performed the experiments: TC DGB MAC LHD SVH EMK KRP AW SZ. Analyzed the data: MUC WH TC TFCM. Wrote the paper: MUC WH TC TFCM.