About the Authors

Shiro Tochitani

tochitani.shiro@gmail.com

Affiliations Division of Development of Mental Functions, Research Center for Child Mental Development, University of Fukui, Fukui 910–1193, Japan, Division of Developmental Higher Brain Functions, Department of Child Development, United Graduate School of Child Development, Osaka University, Kanazawa University, Hamamatsu University School of Medicine, Chiba University and University of Fukui, Osaka 565–0871, Japan, Research and Education Program for Life Science, University of Fukui, Fukui 910–8507, Japan

Takahiro Ikeno

Contributed equally to this work with: Takahiro Ikeno, Tatsuhito Ito, Asuka Sakurai, Tomoki Yamauchi

Affiliation School of Medicine, Faculty of Medical Sciences, University of Fukui, Fukui 910–1193, Japan

Tatsuhito Ito

Contributed equally to this work with: Takahiro Ikeno, Tatsuhito Ito, Asuka Sakurai, Tomoki Yamauchi

Affiliation School of Medicine, Faculty of Medical Sciences, University of Fukui, Fukui 910–1193, Japan

Asuka Sakurai

Contributed equally to this work with: Takahiro Ikeno, Tatsuhito Ito, Asuka Sakurai, Tomoki Yamauchi

Affiliation School of Medicine, Faculty of Medical Sciences, University of Fukui, Fukui 910–1193, Japan

Tomoki Yamauchi

Contributed equally to this work with: Takahiro Ikeno, Tatsuhito Ito, Asuka Sakurai, Tomoki Yamauchi

Affiliation School of Medicine, Faculty of Medical Sciences, University of Fukui, Fukui 910–1193, Japan

Hideo Matsuzaki

Affiliations Division of Development of Mental Functions, Research Center for Child Mental Development, University of Fukui, Fukui 910–1193, Japan, Division of Developmental Higher Brain Functions, Department of Child Development, United Graduate School of Child Development, Osaka University, Kanazawa University, Hamamatsu University School of Medicine, Chiba University and University of Fukui, Osaka 565–0871, Japan, Research and Education Program for Life Science, University of Fukui, Fukui 910–8507, Japan, School of Medicine, Faculty of Medical Sciences, University of Fukui, Fukui 910–1193, Japan

Competing Interests

A patent application has been submitted by the University of Fukui with ST and HM as inventors based on a part of the results described in this paper (Japanese Patent Application No. 2015-017613 Ninshinki-botai tyounai-saikin kakuran ni yoru koudouizyou moderu doubutu no sakusei-houhou to sono youto; the patent application currently has only a Japanese name). There are no further patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ST HM. Performed the experiments: ST T. Ikeno T. Ito AS TY. Analyzed the data: ST. Wrote the paper: ST. Conceived the study: HM.