About the Authors

Hong Ki Min

Contributed equally to this work with: Hong Ki Min, Sung-Min Kim

Affiliations Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea, Division of Rheumatology, Department of Internal Medicine, School of Medicine, The Catholic University of Korea, Seoul, South Korea

Sung-Min Kim

Contributed equally to this work with: Hong Ki Min, Sung-Min Kim

Affiliation Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea

Seung-Ye Baek

Affiliation Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea

Jung-Won Woo

Affiliation Office of Clinical Development, Genexine Inc., Korea Bio Park, Seongnam, Gyeonggi-do, South Korea

Jin-Sil Park

Affiliation Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea

Mi-La Cho

Affiliation Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea

Jennifer Lee

Affiliations Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea, Division of Rheumatology, Department of Internal Medicine, School of Medicine, The Catholic University of Korea, Seoul, South Korea

Seung-Ki Kwok

Affiliations Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea, Division of Rheumatology, Department of Internal Medicine, School of Medicine, The Catholic University of Korea, Seoul, South Korea

Sae Woong Kim

Affiliations Catholic Integrative Medicine Research Institute, College of Medicine, The Catholic University of Korea, Seoul, South Korea, Department of Urology, College of Medicine, The Catholic University of Korea, Seoul, South Korea

Sung-Hwan Park

rapark@catholic.ac.kr

Affiliations Rheumatism Research Center, Catholic Research Institute of Medical Science, The Catholic University of Korea, Seoul, South Korea, Division of Rheumatology, Department of Internal Medicine, School of Medicine, The Catholic University of Korea, Seoul, South Korea

Competing Interests

The authors have declared that no competing interests exist. Genexine Inc. provided support in the form of salaries for author JWW, but did not have any additional role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. The specific roles of these authors are articulated in the "author contributions" section. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SHP SKK HKM. Performed the experiments: SMK SYB JWW JSP. Analyzed the data: HKM MLC JL SKK. Contributed reagents/materials/analysis tools: JL SWK. Wrote the paper: HKM SMK JL SHP.