About the Authors

Simon P. Rout

Affiliation School of Applied Sciences, University of Huddersfield, Huddersfield, UK

Christopher J. Charles

Affiliation School of Applied Sciences, University of Huddersfield, Huddersfield, UK

Charalampos Doulgeris

Affiliation School of Applied Sciences, University of Huddersfield, Huddersfield, UK

Alan J. McCarthy

Affiliation Microbiology Research Group, Institute of Integrative Biology, University of Liverpool, Liverpool, UK

Dave J. Rooks

Affiliation Microbiology Research Group, Institute of Integrative Biology, University of Liverpool, Liverpool, UK

J. Paul Loughnane

Affiliation Microbiology Research Group, Institute of Integrative Biology, University of Liverpool, Liverpool, UK

Andrew P. Laws

Affiliation School of Applied Sciences, University of Huddersfield, Huddersfield, UK

Paul N. Humphreys

p.n.humphreys@hud.ac.uk

Affiliation School of Applied Sciences, University of Huddersfield, Huddersfield, UK

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PNH SPR APL AJM. Performed the experiments: SPC CJC DJR JPL. Analyzed the data: SPR CJC PNH AJM APL CD. Wrote the paper: SPR PNH AJM APL.