About the Authors

Seiji Mori

Contributed equally to this work with: Seiji Mori, Moe Kodaira

Affiliation Department of Molecular Pathology, Osaka University Graduate School of Medicine, Division of Health Sciences, 1–7 Yamada-oka, Suita-shi, Osaka, 565–0871, Japan

Moe Kodaira

Contributed equally to this work with: Seiji Mori, Moe Kodaira

Affiliation Department of Molecular Pathology, Osaka University Graduate School of Medicine, Division of Health Sciences, 1–7 Yamada-oka, Suita-shi, Osaka, 565–0871, Japan

Ayano Ito

Affiliation Department of Molecular Pathology, Osaka University Graduate School of Medicine, Division of Health Sciences, 1–7 Yamada-oka, Suita-shi, Osaka, 565–0871, Japan

Mika Okazaki

Affiliation Department of Molecular Pathology, Osaka University Graduate School of Medicine, Division of Health Sciences, 1–7 Yamada-oka, Suita-shi, Osaka, 565–0871, Japan

Naomasa Kawaguchi

Affiliation Department of Molecular Pathology, Osaka University Graduate School of Medicine, Division of Health Sciences, 1–7 Yamada-oka, Suita-shi, Osaka, 565–0871, Japan

Yoshinosuke Hamada

Affiliation Department of Molecular Pathology, Osaka University Graduate School of Medicine, Division of Health Sciences, 1–7 Yamada-oka, Suita-shi, Osaka, 565–0871, Japan

Yoshikazu Takada

ytakada@ucdavis.edu (YT); matsuura@sahs.med.osaka-u.ac.jp (NM)

Affiliations Departments of Dermatology, Biochemistry and Molecular Medicine, School of Medicine, University of California Davis, Sacramento, California, 95817, United States of America, Graduate Institute of Translational Medicine, College of Medical Science and Technology, Taipei Medical University, 520 Wu-Hsing Street, Taipei, 11031, Taiwan, R.O.C

Nariaki Matsuura

ytakada@ucdavis.edu (YT); matsuura@sahs.med.osaka-u.ac.jp (NM)

Affiliation Department of Molecular Pathology, Osaka University Graduate School of Medicine, Division of Health Sciences, 1–7 Yamada-oka, Suita-shi, Osaka, 565–0871, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SM YT NM. Performed the experiments: SM MK AI MO NK YH. Analyzed the data: SM YT NM. Contributed reagents/materials/analysis tools: YT. Wrote the paper: SM YT.