About the Authors

Margarita Villar

Contributed equally to this work with: Margarita Villar, Nieves Ayllón

Affiliation SaBio, Instituto de Investigación en Recursos Cinegéticos IREC-CSIC-UCLM-JCCM, Ronda de Toledo s/n, 13005, Ciudad Real, Spain

Nieves Ayllón

Contributed equally to this work with: Margarita Villar, Nieves Ayllón

Affiliation SaBio, Instituto de Investigación en Recursos Cinegéticos IREC-CSIC-UCLM-JCCM, Ronda de Toledo s/n, 13005, Ciudad Real, Spain

Katherine M. Kocan

Affiliation Department of Veterinary Pathobiology, Center for Veterinary Health Sciences, Oklahoma State University, Stillwater, Oklahoma, 74078, United States of America

Elena Bonzón-Kulichenko

Affiliation Centro Nacional de Investigaciones Cardiovasculares, Melchor Fernández Almagro 3, 28029, Madrid, Spain

Pilar Alberdi

Affiliation SaBio, Instituto de Investigación en Recursos Cinegéticos IREC-CSIC-UCLM-JCCM, Ronda de Toledo s/n, 13005, Ciudad Real, Spain

Edmour F. Blouin

Affiliation Department of Veterinary Pathobiology, Center for Veterinary Health Sciences, Oklahoma State University, Stillwater, Oklahoma, 74078, United States of America

Sabine Weisheit

Affiliations The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, EH25 9RG, United Kingdom, The Pirbright Institute, Ash Road, Pirbright, Woking, GU24 0NF, United Kingdom

Lourdes Mateos-Hernández

Affiliation SaBio, Instituto de Investigación en Recursos Cinegéticos IREC-CSIC-UCLM-JCCM, Ronda de Toledo s/n, 13005, Ciudad Real, Spain

Alejandro Cabezas-Cruz

Affiliations SaBio, Instituto de Investigación en Recursos Cinegéticos IREC-CSIC-UCLM-JCCM, Ronda de Toledo s/n, 13005, Ciudad Real, Spain, Center for Infection and Immunity of Lille (CIIL), INSERM U1019 – CNRS UMR 8204, Université Lille Nord de France, Institut Pasteur de Lille, Lille, France

Lesley Bell-Sakyi

Affiliations The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, EH25 9RG, United Kingdom, The Pirbright Institute, Ash Road, Pirbright, Woking, GU24 0NF, United Kingdom

Marie Vancová

Affiliations Institute of Parasitology, Biology Centre of the Academy of Sciences of the Czech Republic, Branišovská 31, České Budějovice, CZ-37005, Czech Republic, Faculty of Science, University of South Bohemia, Branišovská 31, České Budějovice, CZ-37005, Czech Republic

Tomáš Bílý

Affiliations Institute of Parasitology, Biology Centre of the Academy of Sciences of the Czech Republic, Branišovská 31, České Budějovice, CZ-37005, Czech Republic, Faculty of Science, University of South Bohemia, Branišovská 31, České Budějovice, CZ-37005, Czech Republic

Damien F. Meyer

Affiliations CIRAD, UMR CMAEE, Site de Duclos, Prise d’eau, F-97170, Petit-Bourg, Guadeloupe, France, INRA, UMR1309 CMAEE, F-34398, Montpellier, France

Jan Sterba

Affiliation Institute of Parasitology, Biology Centre of the Academy of Sciences of the Czech Republic, Branišovská 31, České Budějovice, CZ-37005, Czech Republic

Marinela Contreras

Affiliation SaBio, Instituto de Investigación en Recursos Cinegéticos IREC-CSIC-UCLM-JCCM, Ronda de Toledo s/n, 13005, Ciudad Real, Spain

Nataliia Rudenko

Affiliation Institute of Parasitology, Biology Centre of the Academy of Sciences of the Czech Republic, Branišovská 31, České Budějovice, CZ-37005, Czech Republic

Libor Grubhoffer

Affiliations Institute of Parasitology, Biology Centre of the Academy of Sciences of the Czech Republic, Branišovská 31, České Budějovice, CZ-37005, Czech Republic, Faculty of Science, University of South Bohemia, Branišovská 31, České Budějovice, CZ-37005, Czech Republic

Jesús Vázquez

Affiliation Centro Nacional de Investigaciones Cardiovasculares, Melchor Fernández Almagro 3, 28029, Madrid, Spain

José de la Fuente

jose_delafuente@yahoo.com

Affiliations SaBio, Instituto de Investigación en Recursos Cinegéticos IREC-CSIC-UCLM-JCCM, Ronda de Toledo s/n, 13005, Ciudad Real, Spain, Department of Veterinary Pathobiology, Center for Veterinary Health Sciences, Oklahoma State University, Stillwater, Oklahoma, 74078, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JF M. Villar. Performed the experiments: M. Villar NA KMK EB-K PA EFB SW LB-S AC-C LM-H M. Vancová TB JS MC. Analyzed the data: M. Villar NA KMK PA EB-K JV DFM NR LG JF. Wrote the paper: JF M. Villar NA KMK EB-K JV LB-S.