About the Authors

Kale T. Bentley

bentley2@uw.edu

Affiliation School of Aquatic and Fishery Sciences, Box 355020, University of Washington, Seattle, Washington, United States of America

Daniel E. Schindler

Affiliation School of Aquatic and Fishery Sciences, Box 355020, University of Washington, Seattle, Washington, United States of America

Jonathan B. Armstrong

Affiliations School of Aquatic and Fishery Sciences, Box 355020, University of Washington, Seattle, Washington, United States of America, Wyoming Cooperative Fish and Wildlife Research Unit, Department of Zoology and Physiology, University of Wyoming, Laramie, Wyoming, United States of America

Timothy J. Cline

Affiliation School of Aquatic and Fishery Sciences, Box 355020, University of Washington, Seattle, Washington, United States of America

Gabriel T. Brooks

Affiliation Fish Ecology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KTB DES. Performed the experiments: KTB DES JBA TJC. Analyzed the data: KTB DES JBA TJC. Contributed reagents/materials/analysis tools: KTB DES JBA TJC GTB. Wrote the paper: KTB DES JBA TJC GTB.