About the Authors

Yunda Huang

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Dean Follmann

Affiliation National Institute of Allergy and Infectious Diseases and Biostatistics Research Branch, National Institutes of Health, Bethesda, MD, United States of America

Martha Nason

Affiliation Division of Clinical Research, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, MD, United States of America

Lily Zhang

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Ying Huang

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Devan V. Mehrotra

Affiliation Merck Research Laboratories, North Wales, PA, United States of America

Zoe Moodie

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Barbara Metch

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Holly Janes

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Michael C. Keefer

Affiliation Infectious Disease Division, University of Rochester School of Medicine and Dentistry, Rochester, NY, United States of America

Gavin Churchyard

Affiliation Aurum Institute for Health Research, Johannesburg, South Africa

Merlin L. Robb

Affiliation HJF HIV Program, US Military HIV Research Program, Bethesda, MD, United States of America

Patricia E. Fast

Affiliation Research and Development, International AIDS Vaccine Initiative, New York, New York, United States of America

Ann Duerr

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

M. Juliana McElrath

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Lawrence Corey

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

John R. Mascola

Affiliation Vaccine Research Center, National Institute of Allergy and Infectious Diseases, Bethesda, MD, United States of America

Barney S. Graham

Affiliation Viral Pathogenesis Laboratory, National Institute of Allergy and Infectious Diseases, Bethesda, MD, United States of America

Magdalena E. Sobieszczyk

Affiliation Division of Infectious Diseases, Department of Medicine, Columbia University, New York, New York, United States of America

James G. Kublin

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Michael Robertson

Affiliation Infectious Disease Clinical Research, Merck, Philadelphia, Pennsylvania, United States of America

Scott M. Hammer

Affiliation Division of Infectious Diseases, Department of Medicine, Columbia University, New York, New York, United States of America

Glenda E. Gray

Affiliation University of the Witwatersrand, Johannesburg, South Africa

Susan P. Buchbinder

Affiliation Bridge HIV, San Francisco Department of Public Health, San Francisco, CA, United States of America

Peter B. Gilbert

pgilbert@scharp.org

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Competing Interests

DF, MN, JRM, and BSG are employees of the National Institute of Allergy and Infectious Diseases (NIAID). DVM and MR are employees of Merck. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: Yunda Huang DF MN DVM LC JGK MR SMH GEG SPB PBG. Performed the experiments: Yunda Huang DF LZ MN Ying Huang PBG. Analyzed the data: Yunda Huang DF LZ MN Ying Huang PBG. Wrote the paper: Yunda Huang DF MN LZ Ying Huang DVM ZM BM HJ MCK GC MLR PEF AD MJM LC JRM BSG MES JGK MR SMH GEG SPB PBG. Acquisition of data: Yunda Huang DF BM MLR PBG.