About the Authors

Haolin Liu

Affiliations Howard Hughes Medical Institute, Denver, Colorado, United States of America, Department of Biomedical Research, National Jewish Health, Denver, Colorado, United States of America, Department of Immunology and Microbiology, University of Colorado Denver, Colorado, United States of America

Janice White

Affiliations Department of Biomedical Research, National Jewish Health, Denver, Colorado, United States of America, Department of Immunology and Microbiology, University of Colorado Denver, Colorado, United States of America

Frances Crawford

Affiliations Howard Hughes Medical Institute, Denver, Colorado, United States of America, Department of Biomedical Research, National Jewish Health, Denver, Colorado, United States of America, Department of Immunology and Microbiology, University of Colorado Denver, Colorado, United States of America

Niyun Jin

Affiliations Howard Hughes Medical Institute, Denver, Colorado, United States of America, Department of Biomedical Research, National Jewish Health, Denver, Colorado, United States of America, Department of Immunology and Microbiology, University of Colorado Denver, Colorado, United States of America

Xiangwu Ju

Affiliation State Key Laboratory of Medical Molecular Biology, Institute of Basic Medical Sciences, Chinese Academy of Medical Sciences, Beijing, China

Kangtai Liu

Affiliation State Key Laboratory of Medical Molecular Biology, Institute of Basic Medical Sciences, Chinese Academy of Medical Sciences, Beijing, China

Chengyu Jiang

Affiliation State Key Laboratory of Medical Molecular Biology, Institute of Basic Medical Sciences, Chinese Academy of Medical Sciences, Beijing, China

Philippa Marrack

Affiliations Howard Hughes Medical Institute, Denver, Colorado, United States of America, Department of Biomedical Research, National Jewish Health, Denver, Colorado, United States of America, Department of Immunology and Microbiology, University of Colorado Denver, Colorado, United States of America

Gongyi Zhang

Affiliations Department of Biomedical Research, National Jewish Health, Denver, Colorado, United States of America, Department of Immunology and Microbiology, University of Colorado Denver, Colorado, United States of America

John W. Kappler

kapplerj@njhealth.org

Affiliations Howard Hughes Medical Institute, Denver, Colorado, United States of America, Department of Biomedical Research, National Jewish Health, Denver, Colorado, United States of America, Department of Immunology and Microbiology, University of Colorado Denver, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JK. Performed the experiments: HL JW FC NJ XJ KL. Analyzed the data: HL. Contributed reagents/materials/analysis tools: CJ PM GZ. Wrote the paper: HL JK.