About the Authors

Alexandra T. Boersma

Contributed equally to this work with: Alexandra T. Boersma, Nicholas D. Pyenson

boersma.alex@gmail.com

Affiliations Department of Earth Sciences & Geography, Vassar College, Poughkeepsie, NY, 12604, United States of America, Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, PO Box 37012, Washington, DC, 20013, United States of America

Nicholas D. Pyenson

Contributed equally to this work with: Alexandra T. Boersma, Nicholas D. Pyenson

Affiliations Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, PO Box 37012, Washington, DC, 20013, United States of America, Departments of Mammalogy and Paleontology, Burke Museum of Natural History and Culture, Seattle, WA, 98195, United States of America

Competing Interests

NDP is an academic editor for PLOS ONE. This does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AB NDP. Performed the experiments: AB NDP. Analyzed the data: AB NDP. Contributed reagents/materials/analysis tools: AB NDP. Wrote the paper: AB NDP.