About the Authors

John R. Shorter

john_shorter@ncsu.edu

Affiliations Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America, Program in Genetics, North Carolina State University, Raleigh, North Carolina, United States of America

Matthew Geisz

Affiliation Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, United States of America

Ergi Özsoy

Affiliation Hacettepe University, Department of Biology, Ankara, Turkey

Michael M. Magwire

Current address: Syngenta Biotechnology, Research Triangle Park, North Carolina, United States of America

Affiliation W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America

Mary Anna Carbone

Affiliations Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America, Program in Genetics, North Carolina State University, Raleigh, North Carolina, United States of America

Trudy F. C. Mackay

Affiliations Department of Biological Sciences, North Carolina State University, Raleigh, North Carolina, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America, Program in Genetics, North Carolina State University, Raleigh, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist. MMM is currently employed by a commercial company, Syngenta. However, all of the work done by MMM on the study reported in the manuscript was performed entirely while he was employed by NC State University. Therefore there are no completing interests from a commercial company.

Author Contributions

Conceived and designed the experiments: JRS MG TFCM. Performed the experiments: JRS MG EO MAC. Analyzed the data: JRS MMM. Contributed reagents/materials/analysis tools: TFCM. Wrote the paper: JRS TFCM.