About the Authors

Chengwen Chen

Contributed equally to this work with: Chengwen Chen, Ye Yao

Affiliations State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, People’s Republic of China, Shanghai-MOST Key Laboratory of Health and Disease Genomics, Chinese National Human Genome Center at Shanghai, Shanghai, People's Republic of China, Shanghai Jiao Tong University School of Medicine, Shanghai, People's Republic of China

Ye Yao

Contributed equally to this work with: Chengwen Chen, Ye Yao

Affiliations State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, People’s Republic of China, Center for Computational Systems Biology and School of Mathematical Sciences, Fudan University, Shanghai, People’s Republic of China

Liang Zhang

Affiliation Shanghai-MOST Key Laboratory of Health and Disease Genomics, Chinese National Human Genome Center at Shanghai, Shanghai, People's Republic of China

Minjie Xu

Affiliations State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, People’s Republic of China, Shanghai-MOST Key Laboratory of Health and Disease Genomics, Chinese National Human Genome Center at Shanghai, Shanghai, People's Republic of China

Jianping Jiang

Affiliations State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, People’s Republic of China, Shanghai-MOST Key Laboratory of Health and Disease Genomics, Chinese National Human Genome Center at Shanghai, Shanghai, People's Republic of China

Tonghai Dou

Affiliation State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, People’s Republic of China

Wei Lin

Affiliations State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, People’s Republic of China, Center for Computational Systems Biology and School of Mathematical Sciences, Fudan University, Shanghai, People’s Republic of China

Guoping Zhao

Affiliation Shanghai-MOST Key Laboratory of Health and Disease Genomics, Chinese National Human Genome Center at Shanghai, Shanghai, People's Republic of China

Minren Huang

Affiliation Jiangsu Key Laboratory for Poplar Germplasm Enhancement and Variety Improvement, Nanjing Forestry University, Nanjing, People’s Republic of China

Yan Zhou

zhouy@chgc.sh.cn

Affiliations State Key Laboratory of Genetic Engineering, School of Life Sciences, Fudan University, Shanghai, People’s Republic of China, Shanghai-MOST Key Laboratory of Health and Disease Genomics, Chinese National Human Genome Center at Shanghai, Shanghai, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YZ CC. Performed the experiments: MH TD. Analyzed the data: CC JJ. Contributed reagents/materials/analysis tools: WL YY MX LZ. Wrote the paper: CC YZ GZ.