About the Authors

Vernon Reynolds

vreynolds@btopenworld.com

Affiliations School of Anthropology, Oxford University, Oxford, United Kingdom, Budongo Conservation Field Station, Masindi, Uganda

Andrew W. Lloyd

Affiliation School of Environment & Technology, University of Brighton, Brighton, United Kingdom

Christopher J. English

Affiliation School of Environment & Technology, University of Brighton, Brighton, United Kingdom

Peter Lyons

Affiliation School of Environment & Technology, University of Brighton, Brighton, United Kingdom

Howard Dodd

Affiliation School of Environment & Technology, University of Brighton, Brighton, United Kingdom

Catherine Hobaiter

Affiliations Budongo Conservation Field Station, Masindi, Uganda, School of Psychology, St Andrews University, St Andrews, Scotland

Nicholas Newton-Fisher

Affiliation School of Anthropology and Conservation, University of Kent, Canterbury, United Kingdom

Caroline Mullins

Affiliation School of Psychology, St Andrews University, St Andrews, Scotland

Noemie Lamon

Affiliations Budongo Conservation Field Station, Masindi, Uganda, Institute of Biology, University of Neuchâtel, Neuchâtel, Switzerland

Anne Marijke Schel

Affiliations Budongo Conservation Field Station, Masindi, Uganda, Animal Ecology, Utrecht University, Utrecht, The Netherlands

Brittany Fallon

Affiliations Budongo Conservation Field Station, Masindi, Uganda, School of Psychology, St Andrews University, St Andrews, Scotland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VR AWL. Performed the experiments: CH N-NF CM A-MS NL BF AWL CE PL HD NL. Analyzed the data: AWL VR. Contributed reagents/materials/analysis tools: AWL HD. Wrote the paper: VR AWL CE PL HD CH CM N-NF A-MS NL BF.