About the Authors

Zheng Lao

Affiliations University of Pennsylvania, Perelman School of Medicine, Dept Radiation Oncology, Philadelphia, Pennsylvania, United States of America, Fudan University, Eye & ENT Hospital, Dept Radiation Oncology, Shanghai, China

Catherine J. Kelly

Affiliation Oxford University, Gray Institute for Radiation Oncology, Oxford, United Kingdom

Xiang-Yang Yang

Affiliation University of Pennsylvania, Perelman School of Medicine, Dept Radiation Oncology, Philadelphia, Pennsylvania, United States of America

W. Timothy Jenkins

Affiliation University of Pennsylvania, Perelman School of Medicine, Dept Radiation Oncology, Philadelphia, Pennsylvania, United States of America

Erik Toorens

Affiliation University of Pennsylvania, Perelman School of Medicine, Penn Genomics Analysis Core, Philadelphia, Pennsylvania, United States of America

Tapan Ganguly

Affiliation University of Pennsylvania, Perelman School of Medicine, Penn Genomics Analysis Core, Philadelphia, Pennsylvania, United States of America

Sydney M. Evans

Affiliation University of Pennsylvania, Perelman School of Medicine, Dept Radiation Oncology, Philadelphia, Pennsylvania, United States of America

Cameron J. Koch

kochc@mail.med.upenn.edu

Affiliation University of Pennsylvania, Perelman School of Medicine, Dept Radiation Oncology, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZL C. J. Kelly SME C. J. Koch. Performed the experiments: ZL C. J. Kelly XYY WTJ ET TG SME C. J. Koch. Analyzed the data: ZL C. J. Kelly ET SME C. J. Koch. Contributed reagents/materials/analysis tools: ET TG SME C. J. Koch. Wrote the paper: ZL SME C. J. Koch.