About the Authors

Ryan C. Graebner

Contributed equally to this work with: Ryan C. Graebner, Mitchell Wise

Affiliation Department of Crop and Soil Science, Oregon State University, Corvallis, Oregon, United States of America

Mitchell Wise

Contributed equally to this work with: Ryan C. Graebner, Mitchell Wise

Affiliation Cereal Crops Research, USDA-ARS, Madison, Wisconsin, United States of America

Alfonso Cuesta-Marcos

Affiliation Department of Crop and Soil Science, Oregon State University, Corvallis, Oregon, United States of America

Matthew Geniza

Affiliation Department of Botany and Plant Pathology, Oregon State University, Corvallis, Oregon, United States of America

Tom Blake

Affiliation Plant Sciences and Plant Pathology Department, Montana State University, Bozeman, Montana, United States of America

Victoria C. Blake

Affiliation Crop Improvement and Genetics Research, USDA-ARS, Albany, California, United States of America

Joshua Butler

Affiliation Busch Agricultural Resources, Inc., Fort Collins, Colorado, United States of America

Shiaomen Chao

Affiliation USDA-ARS Cereal Crops Research Unit, Fargo, North Dakota, United States of America

David J. Hole

Affiliation Plants, Soils, and Climate Department, Utah State University, Logan, Utah, United States of America

Rich Horsley

Affiliation Department of Plant Sciences, North Dakota State University, Fargo, North Dakota, United States of America

Pankaj Jaiswal

Affiliation Department of Botany and Plant Pathology, Oregon State University, Corvallis, Oregon, United States of America

Don Obert

Affiliation Limagrain Cereal Seeds, Lafayette, Indiana, United States of America

Kevin P. Smith

Affiliation Department of Agronomy and Plant Genetics, University of Minnesota, St. Paul, Minnesota, United States of America

Steven Ullrich

Affiliation Department of Crop and Soil Science, Washington State University, Pullman, Washington, United States of America

Patrick M. Hayes

patrick.m.hayes@oregonstate.edu

Affiliation Department of Crop and Soil Science, Oregon State University, Corvallis, Oregon, United States of America

Competing Interests

Pankaj Jaiswal is an academic editor for PLOS ONE.

Author Contributions

Conceived and designed the experiments: RCG MW ACM TB VB JB SC DJH RH DO KPS SU PMH PJ MG. Performed the experiments: MW TB VB. Analyzed the data: RCG MG PMH. Contributed reagents/materials/analysis tools: RCG MW ACM TB VB JB SC DJH RH DO KPS SU PMH PJ MG. Wrote the paper: RCG MW MG KPS PMH.