About the Authors

Mai Isosaka

Contributed equally to this work with: Mai Isosaka, Takeshi Niinuma

Affiliation Department of Gastroenterology, Rheumatology and Clinical Immunology, Sapporo Medical University School of Medicine, Sapporo, Japan

Takeshi Niinuma

Contributed equally to this work with: Mai Isosaka, Takeshi Niinuma

Affiliation Department of Molecular Biology, Sapporo Medical University, Sapporo, Japan

Masanori Nojima

Affiliation Center for Translational Research, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Masahiro Kai

Affiliation Department of Molecular Biology, Sapporo Medical University, Sapporo, Japan

Eiichiro Yamamoto

Affiliations Department of Gastroenterology, Rheumatology and Clinical Immunology, Sapporo Medical University School of Medicine, Sapporo, Japan, Department of Molecular Biology, Sapporo Medical University, Sapporo, Japan

Reo Maruyama

Affiliation Department of Gastroenterology, Rheumatology and Clinical Immunology, Sapporo Medical University School of Medicine, Sapporo, Japan

Takayuki Nobuoka

Affiliation Department of Surgery, Surgical Oncology and Science, Sapporo Medical University School of Medicine, Sapporo, Japan

Toshirou Nishida

Affiliation National Cancer Center Hospital East, Kashiwa, Japan

Tatsuo Kanda

Affiliation Department of Surgery, Sanjo General Hospital, Sanjo City, Niigata, Japan

Takahiro Taguchi

Affiliation Division of Human Health and Medical Science, Graduate School of Kuroshio Science, Kochi University, Nankoku, Japan

Tadashi Hasegawa

Affiliation Department of Surgical Pathology, Sapporo Medical University School of Medicine, Sapporo, Japan

Takashi Tokino

Affiliation Medical Genome Science, Research Institute for Frontier Medicine, Sapporo Medical University School of Medicine, Sapporo, Japan

Koichi Hirata

Affiliation Department of Surgery, Surgical Oncology and Science, Sapporo Medical University School of Medicine, Sapporo, Japan

Hiromu Suzuki

hsuzuki@sapmed.ac.jp

Affiliation Department of Molecular Biology, Sapporo Medical University, Sapporo, Japan

Yasuhisa Shinomura

Affiliation Department of Gastroenterology, Rheumatology and Clinical Immunology, Sapporo Medical University School of Medicine, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS YS. Performed the experiments: MI T. Niinuma MK EY. Analyzed the data: T. Niinuma MN RM HS. Contributed reagents/materials/analysis tools: T. Nobuoka T. Nishida TK T. Taguchi TH T. Tokino KH. Wrote the paper: T. Niinuma HS.