About the Authors

Wen-Yi Zhang

Contributed equally to this work with: Wen-Yi Zhang, Li-Ya Wang, Xiu-Shan Zhang, Zhi-Hai Han, Wen-Biao Hu

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Li-Ya Wang

Contributed equally to this work with: Wen-Yi Zhang, Li-Ya Wang, Xiu-Shan Zhang, Zhi-Hai Han, Wen-Biao Hu

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Xiu-Shan Zhang

Contributed equally to this work with: Wen-Yi Zhang, Li-Ya Wang, Xiu-Shan Zhang, Zhi-Hai Han, Wen-Biao Hu

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Zhi-Hai Han

Contributed equally to this work with: Wen-Yi Zhang, Li-Ya Wang, Xiu-Shan Zhang, Zhi-Hai Han, Wen-Biao Hu

Affiliation Navy General Hospital of PLA, Beijing, People’s Republic of China

Wen-Biao Hu

Contributed equally to this work with: Wen-Yi Zhang, Li-Ya Wang, Xiu-Shan Zhang, Zhi-Hai Han, Wen-Biao Hu

Affiliation School of Public Health and Social Work, Queensland University of Technology, Brisbane, Australia

Quan Qian

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Ubydul Haque

Affiliations Emerging Pathogens Institute, University of Florida, Gainesville, Florida, United States of America, Department of Geography, University of Florida, Gainesville, Florida, United States of America

Ricardo J. Soares Magalhaes

Affiliations School of Veterinary Science, The University of Queensland, Brisbane, Australia, Children’s Health and Environment Program, Queensland Children’s Medical Research Institute, The University of Queensland, Brisbane, Australia

Shen-Long Li

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Shi-Lu Tong

Affiliation School of Public Health and Social Work, Queensland University of Technology, Brisbane, Australia

Cheng-Yi Li

licy_60@163.com (C-YL); xxsunhl@163.com (H-LS); sunys1964@hotmail.com (Y-SS)

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Hai-Long Sun

licy_60@163.com (C-YL); xxsunhl@163.com (H-LS); sunys1964@hotmail.com (Y-SS)

Affiliation Institute of Disease Control and Prevention, Academy of Military Medical Science, Beijing, People’s Republic of China

Yan-Song Sun

licy_60@163.com (C-YL); xxsunhl@163.com (H-LS); sunys1964@hotmail.com (Y-SS)

Affiliation Academy of Military Medical Science, Beijing, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WYZ XSZ WBH CYL HLS YSS. Performed the experiments: WYZ LYW SLL. Analyzed the data: WYZ LYW XSZ ZHH CYL HLS YSS. Contributed reagents/materials/analysis tools: CYL HLS YSS. Wrote the paper: WYZ LYW XSZ ZHH WBH QQ UH RJSM SLT CYL HLS YSS.