About the Authors

Tana Wuyun

Affiliation Paulownia Research and Development Center of China, Non-timber Forestry Research and Development of CAF, Weiwu Road, Zhengzhou City 450003, China

Hitomi Amo

Affiliation Food Resources Education and Research Center, Faculty of Agriculture, Kobe University, Hyogo 675–2103, Japan

Jingshi Xu

Affiliation Forestry technology Popularization Station of Beihai City, Beihai 536000, Guang Xi, China

Teng Ma

Affiliation Key Laboratory of Cultivation and Protection for Non-Wood Forest Trees and Non-wood Forest Products of SFA, Central South University of Forestry and Technology, Changsha, Hunan 410004, China

Chiyomi Uematsu

Affiliation Botanical Gardens, Graduate School of Science, Osaka City University, Osaka 576–0004, Japan

Hironori Katayama

hkata@kobe-u.ac.jp

Affiliation Food Resources Education and Research Center, Faculty of Agriculture, Kobe University, Hyogo 675–2103, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HK CU TW. Performed the experiments: TW TM JX HA. Analyzed the data: HA JX. Contributed reagents/materials/analysis tools: TW TM JX HA. Wrote the paper: TW HA HK. Calculated the stastic softwares: HA HK.