About the Authors

Keiko Ikeda

kiikeda@hyo-med.ac.jp

Affiliations Division of Biology, Hyogo College of Medicine, Nishinomiya, Hyogo, Japan, Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Masanori Takahashi

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Shigeru Sato

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Hiroyuki Igarashi

Affiliation Department of Physiology, and Pharmacology, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan

Toru Ishizuka

Affiliation Department of Developmental Biology and Neuroscience, Tohoku University Graduate School of Life Sciences and JST/CREST, Sendai, Miyagi, Japan

Hiromu Yawo

Affiliations Department of Physiology, and Pharmacology, Tohoku University Graduate School of Medicine, Sendai, Miyagi, Japan, Department of Developmental Biology and Neuroscience, Tohoku University Graduate School of Life Sciences and JST/CREST, Sendai, Miyagi, Japan

Satoru Arata

Affiliation Center for Biotechnology, Showa University, Shinagawa, Tokyo, Japan

E. Michelle Southard-Smith

Affiliation Division of Genetic Medicine, Department of Medicine, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Kiyoshi Kawakami

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Hiroshi Onimaru

Affiliation Department of Physiology, Showa University School of Medicine, Shinagawa, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KI HO. Performed the experiments: KI MT HI TI HY EMS KK HO. Analyzed the data: KI SS HO. Contributed reagents/materials/analysis tools: KI HI TI HY SA EMS KK HO. Wrote the paper: KI MES HO.