About the Authors

Changshan Wang

Affiliations Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan, College of Life Sciences, Inner Mongolia University, Hohhot 010021, China

Motohiko Oshima

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan

Goro Sashida

Affiliations Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan, International Research Center for Medical Sciences, Kumamoto University, Kumamoto City 860–0811, Japan

Takahisa Tomioka

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan

Nagisa Hasegawa

Affiliations Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan, Department of Hematology, Chiba University Hospital, Chiba 260–8670, Japan

Makiko Mochizuki-Kashio

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan

Yaeko Nakajima-Takagi

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan

Yoichiro Kusunoki

Affiliation Department of Radiobiology/Molecular Epidemiology, Radiation Effects Research Foundation, Hiroshima 732–0815, Japan

Seishi Kyoizumi

Affiliation Department of Radiobiology/Molecular Epidemiology, Radiation Effects Research Foundation, Hiroshima 732–0815, Japan

Kazue Imai

Affiliation Department of Radiobiology/Molecular Epidemiology, Radiation Effects Research Foundation, Hiroshima 732–0815, Japan

Kei Nakachi

Affiliation Department of Radiobiology/Molecular Epidemiology, Radiation Effects Research Foundation, Hiroshima 732–0815, Japan

Atsushi Iwama

aiwama@faculty.chiba-u.jp

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba 260–8670, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CW YK SK KN AI. Performed the experiments: CW MO GS TT NH MM-K YN-T. Analyzed the data: CW KI AI. Wrote the paper: CW YK SK KN AI.