About the Authors

Kazuya Kusama

Affiliations Department of Endocrine and Neural Pharmacology, Tokyo University of Pharmacy and Life Sciences, 1432–1 Horinouchi, Hachioji, Tokyo, 192–0392, Japan, Laboratory of Theriogenology and Animal Breeding, Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo, Tokyo, 113–8657, Japan

Mikihiro Yoshie

hiro@toyaku.ac.jp (KT); yoshie@toyaku.ac.jp (MY)

Affiliation Department of Endocrine and Neural Pharmacology, Tokyo University of Pharmacy and Life Sciences, 1432–1 Horinouchi, Hachioji, Tokyo, 192–0392, Japan

Kazuhiro Tamura

hiro@toyaku.ac.jp (KT); yoshie@toyaku.ac.jp (MY)

Affiliation Department of Endocrine and Neural Pharmacology, Tokyo University of Pharmacy and Life Sciences, 1432–1 Horinouchi, Hachioji, Tokyo, 192–0392, Japan

Kazuhiko Imakawa

Affiliation Laboratory of Theriogenology and Animal Breeding, Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo, Tokyo, 113–8657, Japan

Keiichi Isaka

Affiliation Department of Obstetrics and Gynecology, Tokyo Medical University, 6-7-1 Nishishinjuku, Tokyo, 160–0023, Japan

Eiichi Tachikawa

Affiliation Department of Endocrine and Neural Pharmacology, Tokyo University of Pharmacy and Life Sciences, 1432–1 Horinouchi, Hachioji, Tokyo, 192–0392, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MY KT. Performed the experiments: KK MY. Analyzed the data: KK MY. Contributed reagents/materials/analysis tools: K. Isaka ET MY KT. Wrote the paper: KT K. Imakawa KK MY.