About the Authors

Nami Suzuki-Hashido

Affiliations Molecular Biology Section, Department of Cellular and Molecular Biology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan, Japan Society for the Promotion of Science, Tokyo, Japan

Takashi Hayakawa

Current Address: Department of Wildlife Science (Nagoya Railroad Co., Ltd.), Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan.; Japan Monkey Centre, Inuyama, Aichi, Japan.

Affiliations Molecular Biology Section, Department of Cellular and Molecular Biology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan, Japan Society for the Promotion of Science, Tokyo, Japan

Atsushi Matsui

Current Address: Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan.

Affiliation Molecular Biology Section, Department of Cellular and Molecular Biology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Yasuhiro Go

Current Address: Center for Novel Science Initiatives, National Institute of Natural Sciences, Okazaki, Aichi, Japan.

Affiliation Molecular Biology Section, Department of Cellular and Molecular Biology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Yoshiro Ishimaru

Affiliation Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Takumi Misaka

Affiliation Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Keiko Abe

Affiliation Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Hirohisa Hirai

Affiliation Molecular Biology Section, Department of Cellular and Molecular Biology, Primate Research Institute, Kyoto University, Inuyama, Aichi, Japan

Yoko Satta

satta@soken.ac.jp (YS); imai.hiroo.5m@kyoto-u.ac.jp (HI)

Affiliation Department of Evolutionary Studies of Biosystems, The Graduate University for Advanced Studies (Sokendai), Hayama, Kanagawa, Japan

Hiroo Imai

satta@soken.ac.jp (YS); imai.hiroo.5m@kyoto-u.ac.jp (HI)

Affiliation Department of Evolutionary Studies of Biosystems, The Graduate University for Advanced Studies (Sokendai), Hayama, Kanagawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NSH TH YI TM YS HI. Performed the experiments: NSH. Analyzed the data: NSH TH AM YG YS. Contributed reagents/materials/analysis tools: YI TM KA HH HI. Wrote the paper: NSH TH HH YS HI.