About the Authors

Fang Chen

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Jing He

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Jianqi Zhang

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Gary K. Chen

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Venetta Thomas

Affiliation Sylvester Comprehensive Cancer Center and Department of Epidemiology and Public Health, University of Miami Miller School of Medicine, Miami, FL, United States of America

Christine B. Ambrosone

Affiliation Department of Cancer Prevention and Control, Roswell Park Cancer Institute, Buffalo, NY, United States of America

Elisa V. Bandera

Affiliation The Cancer Institute of New Jersey, New Brunswick, NJ, United States of America

Sonja I. Berndt

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, MD, United States of America

Leslie Bernstein

Affiliation Division of Cancer Etiology, Department of Population Science, Beckman Research Institute, City of Hope, CA, United States of America

William J. Blot

Affiliations International Epidemiology Institute, Rockville, MD, United States of America, Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt University and the Vanderbilt-Ingram Cancer Center, Nashville, TN, United States of America

Qiuyin Cai

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt University and the Vanderbilt-Ingram Cancer Center, Nashville, TN, United States of America

John Carpten

Affiliation The Translational Genomics Research Institute, Phoenix, AZ, United States of America

Graham Casey

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Stephen J. Chanock

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, MD, United States of America

Iona Cheng

Affiliation Epidemiology Program, Cancer Research Center, University of Hawaii, Honolulu, HI, United States of America

Lisa Chu

Affiliation Cancer Prevention Institute of California, Fremont, CA, United States of America

Sandra L. Deming

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt University and the Vanderbilt-Ingram Cancer Center, Nashville, TN, United States of America

W. Ryan Driver

Affiliation Epidemiology Research Program, American Cancer Society, Atlanta, GA, United States of America

Phyllis Goodman

Affiliation Division of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, WA, United States of America

Richard B. Hayes

Affiliation Division of Epidemiology, Department of Environmental Medicine, New York University Langone Medical Center, New York, NY, United States of America

Anselm J. M. Hennis

Affiliations Chronic Disease Research Centre and Faculty of Medical Sciences, University of the West Indies, Bridgetown, Barbados, Department of Preventive Medicine, Stony Brook University, Stony Brook, NY, United States of America

Ann W. Hsing

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, MD, United States of America

Jennifer J. Hu

Affiliation Sylvester Comprehensive Cancer Center and Department of Epidemiology and Public Health, University of Miami Miller School of Medicine, Miami, FL, United States of America

Sue A. Ingles

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Esther M. John

Affiliations Cancer Prevention Institute of California, Fremont, CA, United States of America, Division of Epidemiology, Department of Health Research & Policy, Stanford University School of Medicine and Stanford Cancer Institute, Stanford, CA, United States of America

Rick A. Kittles

Affiliation Department of Medicine, University of Illinois at Chicago, Chicago, IL, United States of America

Suzanne Kolb

Affiliation Division of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, WA, United States of America

M. Cristina Leske

Affiliation Department of Preventive Medicine, Stony Brook University, Stony Brook, NY, United States of America

Robert C. Millikan

† Deceased.

Affiliation Department of Epidemiology, Gillings School of Global Public Health, and Lineberger Comprehensive Cancer Center, University of North Carolina, Chapel Hill, NC, United States of America

Kristine R. Monroe

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Adam Murphy

Affiliation Department of Urology, Northwestern University, Chicago, IL, United States of America

Barbara Nemesure

Affiliation Department of Preventive Medicine, Stony Brook University, Stony Brook, NY, United States of America

Christine Neslund-Dudas

Affiliation Department of Biostatistics and Research Epidemiology, Henry Ford Hospital, Detroit, MI, United States of America

Sarah Nyante

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, MD, United States of America

Elaine A Ostrander

Affiliation Cancer Genetics Branch, National Human Genome Research Institute, National Institutes of Health, Bethesda, MD, United States of America

Michael F. Press

Affiliation Department of Pathology, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Jorge L. Rodriguez-Gil

Affiliation Sylvester Comprehensive Cancer Center and Department of Epidemiology and Public Health, University of Miami Miller School of Medicine, Miami, FL, United States of America

Ben A. Rybicki

Affiliation Department of Biostatistics and Research Epidemiology, Henry Ford Hospital, Detroit, MI, United States of America

Fredrick Schumacher

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Janet L. Stanford

Affiliation Division of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, WA, United States of America

Lisa B. Signorello

Affiliation Department of Epidemiology, Harvard School of Public Health, Boston, MA, United States of America

Sara S. Strom

Affiliation Department of Epidemiology, The University of Texas M.D. Anderson Cancer Center, Houston, TX, United States of America

Victoria Stevens

Affiliation Epidemiology Research Program, American Cancer Society, Atlanta, GA, United States of America

David Van Den Berg

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Zhaoming Wang

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, MD, United States of America

John S. Witte

Affiliation Institute for Human Genetics, Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, CA, United States of America

Suh-Yuh Wu

Affiliation Department of Preventive Medicine, Stony Brook University, Stony Brook, NY, United States of America

Yuko Yamamura

Affiliation Institute for Human Genetics, Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, CA, United States of America

Wei Zheng

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt University and the Vanderbilt-Ingram Cancer Center, Nashville, TN, United States of America

Regina G. Ziegler

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, MD, United States of America

Alexander H. Stram

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Laurence N. Kolonel

Affiliation Epidemiology Program, Cancer Research Center, University of Hawaii, Honolulu, HI, United States of America

Loïc Le Marchand

Affiliation Epidemiology Program, Cancer Research Center, University of Hawaii, Honolulu, HI, United States of America

Brian E. Henderson

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Christopher A. Haiman

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Daniel O. Stram

stram@usc.edu

Affiliation Department of Preventive Medicine, Keck School of Medicine and Norris Comprehensive Cancer Center, University of Southern California, Los Angeles, CA, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DOS FC CAH BEH. Performed the experiments: FC JH JQZ GKC AHS. Analyzed the data: FC JH JQZ DOS GKC. Contributed reagents/materials/analysis tools: VT CBA EVB SIB LB WJB QC JC GC SJC IC LC SLD WRD PG RBH AJMH AWH JJH SAI EMJ RAK SK MCL RCM KRM AM BN CND SN EAO MFP JLR BAR FS JLS LBS SSS VS DVDB ZW JSW SYW YY WZ RGZ CAH LNK LLM BEH. Wrote the paper: FC DOS.