About the Authors

Nai-Chieh Liu

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

David R. Sargan

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Vicki J. Adams

Affiliation Vet Epi Limited, Birmingham, West Midlands, United Kingdom

Jane F. Ladlow

jfl1001@cam.ac.uk

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Competing Interests

VJA is a self-employed (freelance epidemiologist) of Vet Epi Limited, who was contracted to work on this study and salary was paid by the Kennel Club Charitable Trust. EMMS (Electro-Medical Measurement Systems) provided technical support, to Cambridge Statistics Clinic on use of the plethysmography chambers and their accompanying software. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NCL DRS VJA JFL. Performed the experiments: NCL DRS JFL. Analyzed the data: NCL VJA. Wrote the paper: NCL DRS JFL. Recruited the subjects: NCL DRS VJA JFL.