About the Authors

Zhiyong Zhang

Contributed equally to this work with: Zhiyong Zhang, Hiroko Hatano

Current address: Department of Hemato-Oncology, Institute of Basic Medicine, Shandong Academy of Medical Sciences, Shandong Jinan, Shandong, Peoples Republic of China

Affiliation Nuffield Department of Orthopaedics, Rheumatology and Musculoskeletal Sciences, Botnar Research Centre, University of Oxford, Oxford, United Kingdom

Hiroko Hatano

Contributed equally to this work with: Zhiyong Zhang, Hiroko Hatano

Affiliation Nuffield Department of Orthopaedics, Rheumatology and Musculoskeletal Sciences, Botnar Research Centre, University of Oxford, Oxford, United Kingdom

Jacqueline Shaw

Affiliation Nuffield Department of Orthopaedics, Rheumatology and Musculoskeletal Sciences, Botnar Research Centre, University of Oxford, Oxford, United Kingdom

Marloes Olde Nordkamp

Affiliation RDM Clinical Laboratory Sciences, John Radcliffe Hospital, University of Oxford, Oxford, United Kingdom

Guosheng Jiang

Affiliation Department of Hemato-Oncology, Institute of Basic Medicine, Shandong Academy of Medical Sciences, Shandong Jinan, Shandong, Peoples Republic of China

Demin Li

‡ These authors also contributed equally to this work.

Affiliation RDM Clinical Laboratory Sciences, John Radcliffe Hospital, University of Oxford, Oxford, United Kingdom

Simon Kollnberger

simon.kollnberger@ndorms.ox.ac.uk

‡ These authors also contributed equally to this work.

Affiliation Nuffield Department of Orthopaedics, Rheumatology and Musculoskeletal Sciences, Botnar Research Centre, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZZ HH JS GJ DL SK. Performed the experiments: ZZ HH JS MON SK. Analyzed the data: ZZ HH JS DL SK. Contributed reagents/materials/analysis tools: ZZ JS DL MON SK. Wrote the paper: ZZ HH JS DL SK.