About the Authors

Bernadette Y. Hsu

Contributed equally to this work with: Bernadette Y. Hsu, Yong Ho Bae

Current address: Biomedical Sciences Program, University of California San Francisco, San Francisco, California, United States of America

Affiliations Program in Translational Biomechanics, Institute of Translational Medicine and Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Systems Pharmacology and Translational Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Yong Ho Bae

Contributed equally to this work with: Bernadette Y. Hsu, Yong Ho Bae

yob@mail.med.upenn.edu

Affiliations Program in Translational Biomechanics, Institute of Translational Medicine and Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Systems Pharmacology and Translational Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Keeley L. Mui

Affiliations Program in Translational Biomechanics, Institute of Translational Medicine and Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Systems Pharmacology and Translational Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Shu-Lin Liu

Affiliations Program in Translational Biomechanics, Institute of Translational Medicine and Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Systems Pharmacology and Translational Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Richard K. Assoian

Affiliations Program in Translational Biomechanics, Institute of Translational Medicine and Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Systems Pharmacology and Translational Therapeutics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BYH YHB RKA. Performed the experiments: BYH YHB KLM S-LL. Analyzed the data: BYH YHB. Wrote the paper: BYH YHB KLM RKA.