About the Authors

Hanna Witwicka

Contributed equally to this work with: Hanna Witwicka, Sung-Yong Hwang

Affiliation Department of Cell and Developmental Biology, University of Massachusetts Medical School, Worcester, MA, United States of America

Sung-Yong Hwang

Contributed equally to this work with: Hanna Witwicka, Sung-Yong Hwang

Current address: Center for Drug Evaluation and Research, Food and Drug Administration, Silver Spring, MD, United States of America

Affiliation Department of Cell and Developmental Biology, University of Massachusetts Medical School, Worcester, MA, United States of America

Pablo Reyes-Gutierrez

Affiliation Department of Cell and Developmental Biology, University of Massachusetts Medical School, Worcester, MA, United States of America

Hong Jia

Current address:Department of Developmental Neurobiology, St. Jude Children’s Research Hospital, Memphis, TN, United States of America

Affiliation Department of Cell and Developmental Biology, University of Massachusetts Medical School, Worcester, MA, United States of America

Paul E. Odgren

Affiliation Parallax Pictures, Princeton, MA, United States of America

Leah Rae Donahue

Affiliation The Jackson Laboratory, Bar Harbor, ME, United States of America

Mark J. Birnbaum

Affiliation Department of Biology, Merrimack College, North Andover, MA, United States of America

Paul R. Odgren

paul.odgren@umassmed.edu

Affiliation Department of Cell and Developmental Biology, University of Massachusetts Medical School, Worcester, MA, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SYH HW HJ MJB LRD PRO. Performed the experiments: SYH HW HJ LRD. Analyzed the data: SYH HW PRG PEO HJ LRD MJB PRO. Wrote the paper: SYH HW LRD MJB PRO.