About the Authors

Hua Liu

Affiliation Beijing Key Laboratory of Ornamental Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Laboratory of Urban and rural ecological environment, College of Landscape Architecture, Beijing Forestry University, Beijing, 100083, China

Ming Sun

Affiliation Beijing Key Laboratory of Ornamental Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Laboratory of Urban and rural ecological environment, College of Landscape Architecture, Beijing Forestry University, Beijing, 100083, China

Dongliang Du

Affiliation Beijing Key Laboratory of Ornamental Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Laboratory of Urban and rural ecological environment, College of Landscape Architecture, Beijing Forestry University, Beijing, 100083, China

Huitang Pan

Affiliation Beijing Key Laboratory of Ornamental Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Laboratory of Urban and rural ecological environment, College of Landscape Architecture, Beijing Forestry University, Beijing, 100083, China

Tangren Cheng

Affiliation Beijing Key Laboratory of Ornamental Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Laboratory of Urban and rural ecological environment, College of Landscape Architecture, Beijing Forestry University, Beijing, 100083, China

Jia Wang

Affiliation Beijing Key Laboratory of Ornamental Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Laboratory of Urban and rural ecological environment, College of Landscape Architecture, Beijing Forestry University, Beijing, 100083, China

Qixiang Zhang

zqxbjfu@126.com

Affiliation Beijing Key Laboratory of Ornamental Germplasm Innovation and Molecular Breeding, National Engineering Research Center for Floriculture, Beijing Laboratory of Urban and rural ecological environment, College of Landscape Architecture, Beijing Forestry University, Beijing, 100083, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HL MS QZ. Performed the experiments: HL. Analyzed the data: HL. Contributed reagents/materials/analysis tools: HL DD HP TC JW QZ. Wrote the paper: HL.